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Mollissima in the culmen. The greatest dimorphism is in S. m. dresseri and S. m. sedentaria. 15. Culmen measurements in both sexes in the North East Atlantic. These data show the largestbilled birds in the Baltic/Wadden Sea/North Sea, with declining size in Orkney, Shetland, the Faeroes and Iceland. The smallest culmen sizes are in Faeroes birds. Data from Schiøler (1907, 1914), Heubeck (1993) and personal data. 11. Mean culmen mid-lengths (mm) of S. m. mollissima females from NE Atlantic eider populations.

Given some of the flight distances covered by migrating eiders in a single stage, the Brinkman et al. estimate is considered to be too low. If flight requires about 400kJ per hour (based on Nehls (1996) and Guillemette (1994)), a maximum flight period of about 11 hours is possible. A 10-hour flight at 74kph would equal a range of 740km. Given favourable winds, and energy-efficient formation flying, a single flight exceeding 800km is possible. This equates well with the observational evidence summarised in the movements section (Chapter 3).

There are distinct and persistent gaps in both the central parts of the Russian Arctic and the Canadian Arctic. The distributional gap in Russia (54° longitude) is considerably wider than that in Canada (6° longitude). These gaps can be attributed to the presence of ice all year round, especially landfast sea ice, at these locations. At the highest latitudes, the presence of breeding birds is largely limited by the distribution of ice-free conditions during June. This determines when nesting islands lose their ice bridge connections to the mainland.

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